That's a great question: I'd say its a combination. The microbial building blocks for building the periphyton are still there (just waiting for the summer warm water to come) and I do notice an awful lot of mysterious burrow holes in the dry marl. But there is a seasonal die off, too I would guess, especially during hard dry downs. When that happens getting gambuzia back in the summer water column takes some time, during which mosquitoes can flourish.
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